Ken-ichi Shinoda, Editor-in-Chief, Department of Anthropology Kazuhiro Sakaue, Managing Editor, Department of Anthropology Masanobu Higuchi, Principal Editor, Department of Botany Yoshihito Ohmura, Principal Editor, Department of Botany Toshiaki Kuramochi, Department of Zoology Shoji Hamao, Department of Zoology

Ritsuro Miyawaki, Department of Geology and Paleontology Megumi Saito-Kato, Department of Geology and Paleontology Fumitaka Wakabayashi, Department of Science and Engineering Toshio Kubota, Department of Science and Engineering Yoshikazu Ogawa, Institute for Nature Study Makoto Manabe, Collection Center

National Museum of Nature and Science

Ueno Park, Tokyo 110-8718

Japan

© National Museum of Nature and Science Published on 24 March 2018 Printed by Kyoyu Printing Co., Ltd., Tokyo

ISSN 1881-9109   ISBN 978-4-87803-043-7